5. The Role of Ancient Bees in Shaping Ecosystems

Bees became ever more crucial in forming terrestrial ecosystems as they evolved and diversified. Their effective pollination services let flowering plants spread and diversify quickly, hence changing the makeup of plant populations all around.
The spread of flowering plants made possible by bee pollination had broad consequences for other species. From insects to mammals, new plant species gave food and habitat for a great variety of creatures. Not only bees, but many other new insect lineages were supported in their evolution by the energy-rich nectar and nourishing pollen generated by flowers.
Bees also shaped the way plants reproduce. Many plants evolved systems to use bee behaviour to guarantee outcrossing—mating between genetically different individuals. Certain flowers, for instance, discharge pollen at particular times of day to coincide with the foraging behaviour of their bee pollers. Others have separate male and female stages, therefore lowering the possibility of self-pollination.
Furthermore influencing the evolution of plant defences was the interaction between bees and flowers. Many plants developed traits meant to draw pollinators, but they also had to guard against herbivores. This resulted in the creation of sophisticated chemical defences, many of which are still under research for possible medical use today.
Bees shaped nutrient cycling and soil composition as ecosystem engineers. Their hunting behaviours changed the distribution of plant species over landscapes; their nesting activities aerated the soil and brought organic matter. Bees become important Keystone species in several ecosystems, their presence or absence profoundly influencing the whole community structure.
Additionally indirectly affecting other pollinators was the co-evolution of bees and flowers. Other insects were obliged to adapt or find new niches as blooms become increasingly specialised for bee pollination. This competitive pressure encouraged the development of alternate pollination techniques, therefore adding to the general variety of plant-pollinator interactions observed today.
